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250px-Eosuchus
Holotype E. lerichei skull
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Order: Crocodilia
Superfamily: Gavialoidea
Genus: Eosuchus

Dollo, 1907

Species
  • E. lerichei Dollo, 1907 (type)
  • E. minor (Marsh, 1870 [originally Gavialisminor])

Eosuchus (Dawn Crocodile) is an extinctgenus of gavialoid crocodilian. It is among the most basal of all gavialoids, lying crownward of all other known members of the superfamily, including earlier members such as Thoracosaurusand Eothoracosaurus. Fossils have been found from France as well as easternNorth America in Maryland, Virginia, andNew Jersey. The strata from which specimens have been found date back to the late Paleocene and early Eoceneepochs.

Species Edit

220px-Eosuchus jaw

Lower jaw of the holotype

The name Eosuchus was first used in 1907 to describe a single specimen found from northern France near theBelgian border, assigned to the type species E. lerichei.[1] A second species,E. minor, was named in 2006 from material found from the United States, having been referred in the past to several different genera of crocodilians.E. minor differs from E. lerichei on the basis of a noticeably wider nasal andprefrontals positioned anteriorly further up the skull than the lacrimals

A new species of GavialisG. minor, was named by Othniel Charles Marsh in 1870 on the basis of the holotypespecimen YPM 282, consisting of cranial fragments and isolated vertebrae found from the Manasquan Formation in Monmouth County, New Jersey. The locality dates back to theYpresian stage of the early Eocene. The species name refers to the relatively small estimated size of the animal, estimated at no more than 2 meters, when compared to other gavialoids such as the modern gharial, which can grow up to 5 meters in length.

However, this species was later recognized as distinct from Gavialis on the basis of certain aspects of the known cranial material, in particular the large foramen aerum of the quadrateformed from the epithelial tube that connects the pneumatic chambers of the quadrate and articular. Another diagnostic feature thought to distinguish the species from Gavialiswas the narrow interfenestral bar of theparietal bone that is relatively smooth and unsculptured when compared to other gavialoids such as Thoracosaurus. The name Thecachampsoides was proposed for the species G. minor in 1986.[2] A close relationship between T. minor and Eosuchus lerichei was always evident, yet it was not until 2006 that the name Eosuchus was applied to the T. minor specimens, specifically on the basis of a fairly complete specimen called NJSM 15437 from the Vincetown Formation in New Jersey, of which there is a visibly exposed braincase which aids greatly in the taxonomic classification of the genus. The examination of specimens of T. minorwith those of E. lerichei yielded many similarities between the two species, including the foramen aerum as well as other features such as a long nasal process between the premaxillae,dentary alvioli arranged in pairs, and a W-shaped basioccipital tuberosity.[3]

Other material present from the Aquia Formation of Maryland and Virginia, which dates back to the earlyPaleocene, tends to be more complete. Some specimens found from these localities are known from nearly complete skulls that provide a more detailed view of the phylogenetic position of Eosuchus, and further aid in distinguishing E. minor from other gavialoids.

Historically, many other species of extinct gavialoids have been assigned to the genus Gavialis (of which the modern gharial, G. gangeticus, is a member of). These include forms such as Thoracosaurus.

Phylogenetics Edit

Eogavialis is often referred to as a "thoracosaur", a group that was constructed to include many basal forms of gavialoids that cannot be placed into any specific families of their own, yet do not form a true clade. The genus is distinct from the laterEogavialis and Neogene gavialoids in still having a thin descending laminapositioned next to the pterygoid.

A phylogenetic analysis conducted in 1996 suggested that Eosuchus may be closely related to tomistomines such as the extant Tomistoma. There is further evidence of this relationship shown in the humeri of these crocodilians; bothEosuchus and tomistomines have a deeply concave deltopectoral crest, unlike the more triangular crests ofGavialis. A nasal-premaxilla contact and similar dentary and maxillary tooth counts also seem to suggest that there may be a close relationship between the genus and later tomistomines, although these features represent more primitive conditions that changed later in gavialoid history. Despite this, it is currently accepted that a close relationship between Eosuchus and tomistomines is not the case, and that the similarities between the two may just be superficial and that these characteristics are plesiomorphic to all gavialoids, being lost in more derived members such as the modern Gavialis.

Paleobiology Edit

220px-Eosuchus osteoderms-1

Osteoderms

The strata from which both species ofEosuchus have been found were though to have formed in a marginal marinedepositional environment, and thus probably reflect the actual environments that these animals would have inhabited. It has been proposed that early gavialoids were originally salt-tolerant coastal forms,[4] and the evidence seen in the case of Eosuchusis consistent with this theory. One specimen of E. minor from the Aquia Formation, USNM 299730, has a fossiloyster attached to the dorsal surface of the rostrum.

The fact that the two species ofEosuchus lived on either side of theAtlantic Ocean implies that these populations may have been separated geographically from one another while not necessarily having to be separated stratigraphically (that is, if the temporal ranges of the two species coincide with one another). More importantly, the separate biogeographic ranges of the two species may be evidence for a transoceanic dispersal event from one continent to the other. Since the presumed ages of the localities from which specimens have been found are quite similar yet inexact, it is currently unknown just what continent this dispersal event may have originated. A recent reevaluation of the holotype material of E. lerichei, which in the past has been poorly studied, suggests that it is the more basal species and thus would have been the ancestor of E. minor in Europe.[5]

Other taxa referred to Eosuchus Edit

A Triassic archosauromorph was originally named Eosuchus, although it is completely unrelated to the gavialoid of the same name. The name of the archosaur was later changed toNoteosuchus due to this preoccupation.

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